Neuroscience

Hedco Neurosciences Building 120
(213) 740-6090
FAX: (213) 740-2534
Email: neuroscienceundergraduate@dornsife.usc.edu
usc.edu/programs/neuroscience

Director: Pat Levitt, Ph.D.

Participating Faculty: See Biological Sciences, Computer Science, Biomedical Engineering, Philosophy, Psychology, Engineering, Gerontology, Medicine and Pharmacy in this catalogue.

The requirements for this program will change beginning fall 2013.

Show revised version

Bachelor of Arts in Neuroscience

Co-directors: Sarah Bottjer, Ph.D., and Irving Biederman, Ph.D.

Undergraduate Advisers: Lauren Evashenk, evashenk@dornsife.usc.edu, and Jessika Hulewsky, jbonds@dornsife.usc.edu

Grade Requirements

A grade of C- or higher is required to count toward major requirements.

Core Requirements (32 units) Units
BISC 220L General Biology: Cell Biology and Physiology 4
BISC 421 Neurobiology 4
CHEM 103Lx General Chemistry for the Environment and Life, or
CHEM 105aL General Chemistry 4
MATH 125 Calculus I 4
NEUR 408 Systems Neuroscience: From Synapses to Perception 4
PSYC 100 Introduction to Psychology 4
PSYC 274L* Statistics 4
PSYC 440 Introduction to Cognitive Neuroscience 4

*An equivalent course may be substituted with permission.

(8 core + 4* or 5* elective: 48 units)

Four or five upper-division elective courses (minimum 16 units) from the following list are required. At least one course in the upper-division electives must carry a lab (“L”) designation or be NEUR 490.

Electives (16 units) Units
ANTH 306 Primate Social Behavior 4
BISC 307L General Physiology 4
BISC 313 Evolution and Population Genetics 4
BISC 320L Molecular Biology 4
BISC 325 Genetics 4
BISC 330L Biochemistry 4
BISC 403 Advanced Molecular Biology 4
BISC 406L Biotechnology 4
BISC 410 Applications of Molecular Biology to Medicine 4
BISC 411 Cell Biology 4
BISC 422L Neurobiology Laboratory 4
BISC 423 Epilepsy to Ecstasy: Biological Basis of Neurological Disorders 4
BISC 424 Brain Architecture 4
BISC 426 Principles of Neural Development 4
BISC 462 Seminar in Neurobiology 2, max 4
BISC 480 Developmental Biology 4
BISC 490x Direct Research 4
BME 402 Control and Communication in the Nervous System 3
CSCI 460 Introduction to Artificial Intelligence 3
GERO 414 Neurobiology of Aging 4
GERO 415 Neuroaffective Disorders of Aging 4
MATH 265 Mathematical and Computational Methods for Neuroscience 4
PSYC 301L Cognitive Processes 4
PSYC 304L Sensation and Perception 4
PSYC 305 Learning and Memory 4
PSYC 320 Principles of Psychobiology 4
PSYC 326 Behavioral Neuroscience 4
PSYC 339L Origins of the Mind 4
PSYC 404L Psychophysiology of Emotion 4
PSYC 420 Animal Behavior 4
PSYC 425 Functional Imaging of the Human Brain 4
PSYC 426 Motivated Behavior 4
PSYC 438 Behavioral Genetics 4
PSYC 450L Neural Network Models of Social and Cognitive Processes 4

A graduate class from the following list can be substituted for an upper division elective with permission.

BME 575L Computational Neuroengineering 3
CSCI 564 Brain Theory and Artificial Intelligence 3
NEUR 524* Advanced Overview of Neuroscience 4
NEUR 531* Molecular and Cellular Neurobiology 4
NEUR 532* Systems and Behavioral Neurobiology 3
PSYC 506* Learning and Cognition 4
PSYC 510* Visual Cognition 4
PSYC 540* Cognitive Neuroscience 4
PSYC 544* Psychophysiology 4
PSYC 545* Neuropsychology 4
PSYC 547* Functional Neuroanatomy 4
PSYC 551* Decision Neuroscience 4

*With minimum GPA of 3.3 and permission of co-directors and instructor

Bachelor of Arts in Neuroscience

Co-directors: Sarah Bottjer, Ph.D., and Irving Biederman, Ph.D.

Undergraduate Advisers: Lauren Evashenk, evashenk@dornsife.usc.edu, and Jessika Hulewsky, jbonds@dornsife.usc.edu

Grade Requirements

A grade of C- or higher is required to count toward major requirements.

Core Requirements (32 units) Units
BISC 220L General Biology: Cell Biology and Physiology, or
BISC 221L Advanced General Biology: Cell Biology and Physiology 4
BISC 421 Neurobiology 4
CHEM 103Lx General Chemistry for the Environment and Life, or
CHEM 105aL General Chemistry 4
MATH 125 Calculus I 4
NEUR 408 Systems Neuroscience: From Synapses to Perception 4
PSYC 100 Introduction to Psychology 4
PSYC 274L* Statistics 4
PSYC 440 Introduction to Cognitive Neuroscience 4

*An equivalent course may be substituted with permission.

(8 core + 4* or 5* elective: 48 units)

Four or five upper-division elective courses (minimum 16 units) from the following list are required. At least one course in the upper-division electives must carry a lab (“L”) designation or be NEUR 490. No more than 4 units of NEUR 490x may be used to fulfill the upper-division elective requirement.

Electives (16 units) Units
ANTH 306 Primate Social Behavior 4
BISC 307L General Physiology 4
BISC 313 Evolution and Population Genetics 4
BISC 320L Molecular Biology 4
BISC 325 Genetics 4
BISC 330L Biochemistry 4
BISC 403 Advanced Molecular Biology 4
BISC 406L Biotechnology 4
BISC 410 Applications of Molecular Biology to Medicine 4
BISC 411 Cell Biology 4
BISC 422L Neurobiology Laboratory 4
BISC 423 Epilepsy to Ecstasy: Biological Basis of Neurological Disorders 4
BISC 424 Brain Architecture 4
BISC 426 Principles of Neural Development 4
BISC 462 Seminar in Neurobiology 2, max 4
BISC 480 Developmental Biology 4
BME 402 Control and Communication in the Nervous System 3
CSCI 460 Introduction to Artificial Intelligence 3
GERO 414 Neurobiology of Aging 4
GERO 415 Neuroaffective Disorders of Aging 4
MATH 265 Mathematical and Computational Methods for Neuroscience 4
NEUR 490x Direct Research 2, max 4
PSYC 301L Cognitive Processes 4
PSYC 304L Sensation and Perception 4
PSYC 305 Learning and Memory 4
PSYC 320 Principles of Psychobiology 4
PSYC 326 Behavioral Neuroscience 4
PSYC 339L Origins of the Mind 4
PSYC 404L Psychophysiology of Emotion 4
PSYC 420 Animal Behavior 4
PSYC 424 Neuropsychology 4
PSYC 425 Functional Imaging of the Human Brain 4
PSYC 426 Motivated Behavior 4
PSYC 438 Behavioral Genetics 4
PSYC 450L Neural Network Models of Social and Cognitive Processes 4

A graduate class from the following list can be substituted for an upper division elective with permission.

BME 575L Computational Neuroengineering 3
CSCI 564 Brain Theory and Artificial Intelligence 3
NEUR 524* Advanced Overview of Neuroscience 4
NEUR 531* Molecular and Cellular Neurobiology 4
NEUR 532* Systems and Behavioral Neurobiology 3
PSYC 506* Learning and Cognition 4
PSYC 510* Visual Cognition 4
PSYC 540* Cognitive Neuroscience 4
PSYC 544* Psychophysiology 4
PSYC 545* Neuropsychology 4
PSYC 547* Functional Neuroanatomy 4
PSYC 551* Decision Neuroscience 4

*With minimum GPA of 3.3 and permission of co-directors and instructor

The requirements for this program will change beginning fall 2013.

Show revised version

Bachelor of Science in Neuroscience

Co-directors: Sarah Bottjer, Ph.D., and Irving Biederman, Ph.D.

Undergraduate Advisers: Lauren Evashenk, evashenk@dornsife.usc.edu and Jessika Hulewsky, jbonds@dornsife.usc.edu

Grade Requirements

A grade of C- or higher is required to count toward major requirements.

Core Requirements (56 units) Units
BISC 220L General Biology: Cell Biology and Physiology 4
BISC 421 Neurobiology 4
CHEM 105abL General Chemistry 4-4
CHEM 322abL Organic Chemistry 4-4
MATH 125 Calculus I 4
MATH 265 Mathematical and Computational Methods for Neuroscience 4
NEUR 408 Systems Neuroscience: From Synapses to Perception 4
PHYS 135abL Physics for the Life Sciences 4-4
PSYC 100 Introduction to Psychology 4
PSYC 274* Statistics I 4
PSYC 440 Introduction to Cognitive Neuroscience 4

*An equivalent course may be substituted with permission.

(14 core + 5* or 6* elective: 76 units)

Five or six upper-division elective courses (minimum 20 units) from the following list are required. At least one course in the upper-division electives must carry a lab (“L”) designation or be 490.

Electives (20 units) Units
ANTH 306 Primate Social Behavior 4
BISC 307L General Physiology 4
BISC 313 Evolution and Population Genetics 4
BISC 320L Molecular Biology 4
BISC 325 Genetics 4
BISC 330L Biochemistry 4
BISC 403 Advanced Molecular Biology 4
BISC 406L Biotechnology 4
BISC 410 Applications of Molecular Biology to Medicine 4
BISC 411 Cell Biology 4
BISC 422L Neurobiology Laboratory 4
BISC 423 Epilepsy to Ecstasy: Biological Basis of Neurological Disorders 4
BISC 424 Brain Architecture 4
BISC 426 Principles of Neural Development 4
BISC 462 Seminar in Neurobiology 2, max 4
BISC 480 Developmental Biology 4
BISC 490x Directed Research 4
BME 402 Control and Communication in the Nervous System 3
CSCI 460 Introduction to Artificial Intelligence 3
GERO 414 Neurobiology of Aging 4
GERO 415 Neuroaffective Disorders of Aging 4
PSYC 301L Cognitive Processes 4
PSYC 304L Sensation and Perception 4
PSYC 305 Learning and Memory 4
PSYC 320 Principles of Psychobiology 4
PSYC 326 Behavioral Neuroscience 4
PSYC 339L Origins of the Mind 4
PSYC 404L Psychophysiology of Emotion 4
PSYC 420 Animal Behavior 4
PSYC 425 Functional Imaging of the Human Brain 4
PSYC 426 Motivated Behaviors 4
PSYC 438 Behavioral Genetics 4
PSYC 450L Neural Network Models of Social and Cognitive Processes 4
PSYC 490x Directed Research 4

A graduate class from the following list can be substituted for an upper-division elective with permission.

BME 575L* Computational Neuroengineering 3
CSCI 564* Brain Theory and Artificial Intelligence 3
NEUR 524* Advanced Overview of Neuroscience 4
NEUR 531* Molecular and Cellular Neurobiology 4
NEUR 532* Systems and Behavioral Neurobiology 3
PSYC 506* Learning and Cognition 4
PSYC 510* Visual Cognition 4
PSYC 540* Cognitive Neuroscience 4
PSYC 544* Psychophysiology 4
PSYC 545* Neuropsychology 4
PSYC 547* Functional Neuroanatomy 4
PSYC 551* Decision Neuroscience 4

*With minimum GPA of 3.3 and permission of co-directors and instructor

Bachelor of Science in Neuroscience

Co-directors: Sarah Bottjer, Ph.D., and Irving Biederman, Ph.D.

Undergraduate Advisers: Lauren Evashenk, evashenk@dornsife.usc.edu and Jessika Hulewsky, jbonds@dornsife.usc.edu

Grade Requirements

A grade of C- or higher is required to count toward major requirements.

Core Requirements (56 units) Units
BISC 220L General Biology: Cell Biology and Physiology, or
BISC 221L Advanced General Biology: Cell Biology and Physiology 4
BISC 421 Neurobiology 4
CHEM 105aL General Chemistry, or
CHEM 115aL Advanced General Chemistry 4
CHEM 105bL General Chemistry, or
CHEM 115bL Advanced General Chemistry 4
CHEM 322abL Organic Chemistry 4-4
MATH 125 Calculus I 4
MATH 265 Mathematical and Computational Methods for Neuroscience 4
NEUR 408 Systems Neuroscience: From Synapses to Perception 4
PHYS 135aL Physics for the Life Sciences, or
PHYS 151 Fundamentals of Physics I: Mechanics and Thermodynamics 4
PHYS 135bL Physics for the Life Sciences, or
PHYS 152 Fundamentals of Physics II: Electricity and Magnetism 4
PSYC 100 Introduction to Psychology 4
PSYC 274* Statistics I 4
PSYC 440 Introduction to Cognitive Neuroscience 4

*An equivalent course may be substituted with permission.

(14 core + 5* or 6* elective: 76 units)

Five or six upper-division elective courses (minimum 20 units) from the following list are required. At least one course in the upper-division electives must carry a lab (“L”) designation or be 490. No more than 4 units of NEUR 490x may be used to fulfill the upper-division elective requirement.

Electives (20 units) Units
ANTH 306 Primate Social Behavior 4
BISC 307L General Physiology 4
BISC 313 Evolution and Population Genetics 4
BISC 320L Molecular Biology 4
BISC 325 Genetics 4
BISC 330L Biochemistry 4
BISC 403 Advanced Molecular Biology 4
BISC 406L Biotechnology 4
BISC 410 Applications of Molecular Biology to Medicine 4
BISC 411 Cell Biology 4
BISC 422L Neurobiology Laboratory 4
BISC 423 Epilepsy to Ecstasy: Biological Basis of Neurological Disorders 4
BISC 424 Brain Architecture 4
BISC 426 Principles of Neural Development 4
BISC 462 Seminar in Neurobiology 2, max 4
BISC 480 Developmental Biology 4
BME 402 Control and Communication in the Nervous System 3
CSCI 460 Introduction to Artificial Intelligence 3
GERO 414 Neurobiology of Aging 4
GERO 415 Neuroaffective Disorders of Aging 4
NEUR 490x Directed Research 2, max 4
PSYC 301L Cognitive Processes 4
PSYC 304L Sensation and Perception 4
PSYC 305 Learning and Memory 4
PSYC 320 Principles of Psychobiology 4
PSYC 326 Behavioral Neuroscience 4
PSYC 339L Origins of the Mind 4
PSYC 404L Psychophysiology of Emotion 4
PSYC 420 Animal Behavior 4
PSYC 424 Neuropsychology 4
PSYC 425 Functional Imaging of the Human Brain 4
PSYC 426 Motivated Behaviors 4
PSYC 438 Behavioral Genetics 4

A graduate class from the following list can be substituted for an upper-division elective with permission.

BME 575L* Computational Neuroengineering 3
CSCI 564* Brain Theory and Artificial Intelligence 3
NEUR 524* Advanced Overview of Neuroscience 4
NEUR 531* Molecular and Cellular Neurobiology 4
NEUR 532* Systems and Behavioral Neurobiology 3
PSYC 506* Learning and Cognition 4
PSYC 510* Visual Cognition 4
PSYC 540* Cognitive Neuroscience 4
PSYC 544* Psychophysiology 4
PSYC 545* Neuropsychology 4
PSYC 547* Functional Neuroanatomy 4
PSYC 551* Decision Neuroscience 4

*With minimum GPA of 3.3 and permission of co-directors and instructor

The requirements for this program will change beginning fall 2013.

Show revised version

Honors Program in Neuroscience

An honors program is available to outstanding students already pursuing a B.A. degree in Neuroscience. This program offers students an opportunity to participate in undergraduate research, experience in writing an honors thesis summarizing the completed research and experience in an honors seminar. Honors students must register for NEUR 490x Directed Research, or an equivalent course in another department. NEUR 490x can replace one of the courses from either of the two lists above. Honors students are also required to take one semester of NEUR 494x or an equivalent course in addition to fulfilling all the requirements for the B.A. degree. Approval from the program in neuroscience is required before credit will be given for either of these two research-related courses in another department. The student earning honors in neuroscience must have a minimum overall GPA of 3.5 at graduation. This program leads to the designation on the transcript of Bachelor of Arts in Neuroscience with Honors.

Honors Program in Neuroscience

An honors program is available to outstanding students who are pursuing a B.A. or B.S. degree in Neuroscience. This program offers students exceptional opportunities to participate in undergraduate research, culminating in the experience of writing an honors thesis summarizing their completed research. Honors students must register for 4 units of Directed Research (NEUR 490x). Honors students are also required to take two semesters of the Honors Seminar (BISC 493x as one of their upper-division electives, 1 unit/semester), in which small groups of students discuss recent findings in neuroscience literature and their own research. After completing the honors seminar, honors students also take one semester of honors thesis (BISC 494x, 2 units), in which students write their senior thesis. Students earning honors in neuroscience must have a minimum overall GPA of 3.5 at graduation. This program leads to the designation on the transcript of Bachelor of Arts/Science in Neuroscience with Honors.

Minor in Neuroscience

Co-coordinators: Sarah Bottjer, Ph.D., and Irving Biederman, Ph.D.

Undergraduate Advisers: Lauren Evashenk, evashenk@dornsife.usc.edu, and Jessika Hulewsky, jbonds@dornsife.usc.edu

Grade Requirements

A grade of C- or higher is required to count toward minor requirements.

Core Requirements (20 units) Units
PSYC 274* Statistics I 4
BISC 421** Neurobiology 4
NEUR 408** Systems Neuroscience: From Synapses to Perception 4
PSYC 440*** Introduction to Cognitive Neuroscience 4
Electives A 300- or 400-level course from the elective list for majors 4

*An equivalent course may be substituted with permission

**Prerequisite: BISC 220 or BISC 221

***Prerequisite: PSYC 100

Recommended but not required: CHEM 103Lx (or CHEM 105La) plus MATH 125.

Students who have not already taken the prerequisites will need to take a total of 28 units to satisfy the requirements of a minor in neuroscience. The range of units will vary from 20 to 28 depending on a student’s background.

Master of Science in Neuroscience

Coordinator: Pat Levitt, Ph.D.

Enrollment of graduate students as master’s degree candidates is not encouraged and is reserved for special, terminal circumstances. To satisfy the requirements for the M.S. degree the student must take all the course work required of Ph.D. students for a minimum of 24 units. Completion of the degree requires the submission of a short, formal paper of original research that is approved by three members of the neuroscience graduate program faculty. Students must also satisfy residency and other requirements of the Graduate School.

Doctor of Philosophy in Neuroscience

Coordinator: Pat Levitt, Ph.D.

Application deadline: December 15

Breadth of interests and training are major features of the graduate program in neuroscience. Wide and varied skills in many research areas characterize the faculty of the program. Close contact between faculty and students is considered of major importance in this highly interdisciplinary field.

Training is given in several areas of specialization: behavioral and systems neuroscience, cellular and molecular neurobiology, cognitive neuroscience, computational neuroscience, neuroengineering and neuroscience of aging and development.

Applicants should normally have defined an interest in one or two specializations. A final choice of the specialization will be made during the first year.

Admission Requirements

A baccalaureate degree in a field relevant to the student’s graduate goals is required.

Appropriate fields would include neuroscience, biology, chemistry, computer science, linguistics, psychology and many areas of engineering. Undergraduate study should provide evidence of proficiency in mathematics, including statistics. Students planning to enter the specialization in computational and mathematical neuroscience should have taken course work in calculus and, where possible, linear algebra and computer programming. Applicants who are accepted with minor deficiencies are expected to correct these during the first year.

Applications require forms from both the university and the program. These may be obtained from: Coordinator, Graduate Program in Neuroscience, University of Southern California, Los Angeles, CA 90089-2520.

Degree Requirements

These degrees are awarded under the jurisdiction of the Graduate School. Refer to the Requirements for Graduation and the Graduate School for general regulations. All courses applied toward the degrees must be courses accepted by the Graduate School.

Advisory Committee

The student will be advised during the first year by the Graduate Affairs Committee. As soon as the student has selected a specialization, an Advisory Committee of appropriate faculty will be appointed. This committee will be chaired by the thesis adviser, when chosen. The purpose of the Advisory Committee is to help the student in the selection of courses and research; to monitor the student’s progress; to insure preparation for the qualifying examination; and to administer that examination.

Course Requirements

A minimum of 60 units is required, consisting of formal courses, seminars and research credits. At least 24 of the 60 units are to be formal graduate course work (lecture or seminar courses). During the first year the student is expected to complete the core courses in neuroscience (NEUR 524), one key course, NEUR 538 Neuroscience Ethics and Professionalization, and two semesters of NEUR 539. Other courses in the area of specialization may also be taken in the first year and will be taken in subsequent years.

Core Course: NEUR 524 Advanced Overview of Neuroscience (4 units), will be taken by all students in the fall of their first year to provide an integrated multilevel view of neuroscience. To take the core course, students should have mastered the material currently taught in BISC 421. (Students will be expected to review a detailed syllabus and reading list for BISC 421 to identify their level of knowledge prior to their arrival at USC and will receive advice at Orientation on whether to take BISC 421 or read recommended material to remedy their deficiencies.)

Key Courses: All students will be required to complement their thesis-directed studies with a “breadth with depth” requirement by taking three key courses, one each from three of the four tracks listed below. Each key course will be for 3 or 4 units. (At least one of these courses will serve to advance thesis-related study as well.)

Cellular, Molecular and Developmental Neuroscience Track units
NEUR 531 Molecular and Cellular Neurobiology 4
BISC 426 Principles of Neural Development 4
Cognitive Neuroscience Track units
PSYC 540 Cognitive Neuroscience 4
Computational Neuroscience and Neuroengineering Track units
BME 575L Computational Neuroengineering 3
NEUR 535 Brain Theory and Artificial Intelligence 3
Systems and Behavioral Neuroscience Track units
NEUR 532 Systems and Behavioral Neurobiology 3

All students are required to take NEUR 538 Neuroscience Ethics and Professionalization (1 unit).

It is required that all neuroscience Ph.D. students demonstrate competence in statistics in fulfillment of their Ph.D. requirements.

Qualifying Examination

The qualifying examination concentrates on the student’s ability to demonstrate a grasp of the major area of interest chosen and its relation to other areas of training offered in the program. The examination is partly written and partly oral and is designed to test the student’s ability to meet the demands of the profession.

Dissertation

An acceptable dissertation based on completion of an original investigation is required. The candidate must defend an approved draft of the dissertation in an oral examination.

Courses of Instruction

Neuroscience (NEUR)

The terms indicated are expected but are not guaranteed. For the courses offered during any given term, consult the Schedule of Classes.

NEUR 390 Special Problems (1-4) Supervised, individual studies. No more than one registration permitted. Enrollment by petition only.

NEUR 408 Systems Neuroscience: From Synapses to Perception (4, Sp) Sensory systems to illustrate basic concepts regarding the functional organization of the brain, from the microscopic arrangement of neural circuits to global processes such as perception. Prerequisite: BISC 220 or BISC 221.

NEUR 426 Principles of Neural Development (4, Sp) (Enroll in BISC 426)

NEUR 490x Directed Research (2-4, max 8, FaSpSm) Individual research and readings. Not available for graduate credit.

NEUR 494x Honors Thesis (2, FaSp) Not available for graduate credit. Programmatic approval.

NEUR 524 Advanced Overview of Neuroscience (4, Fa) Study of the nervous system at multiple levels through the analysis of four themes: motor control; emotion, motivation, and decision-making; memory and learning; and vision. Open only to graduate students. Prerequisite: BISC 421.

NEUR 531 Molecular and Cellular Neurobiology (4, Fa) Introduces fundamental principles of advanced molecular and cellular neurobiology including proteins and nucleic acids, cell biology of neurons and glia, synaptic transmission and neuronal signaling. Open only to graduate students.

NEUR 532 Systems and Behavioral Neurobiology (3, Fa) Systems and behavioral neurobiology: hierarchical mechanisms controlling behavior, experimental techniques; perceptual (visual, auditory, somatosensory) systems; sensorimotor systems; motivated behavior; learning, memory and adaptation. Open only to graduate students. Prerequisite: NEUR 524.

NEUR 533 Cognitive Neuroscience (4, Sp) (Enroll in PSYC 540)

NEUR 534L Computational Neuroengineering (3) (Enroll in BME 575L)

NEUR 535 Brain Theory and Artificial Intelligence (3) (Enroll in CSCI 564)

NEUR 538 Neuroscience Ethics and Professionalization (1, Fa) Exposes students to ethical issues in scientific research, especially for neuroscience; scientific integrity and professional roles for the academician and neuroscientist. Open only to graduate students.

NEUR 539 Seminar in Neurobiology (1, max 4, FaSp) (Duplicates credit in former BISC 539.)

NEUR 541 Neurobiology of Disease (3, Sp) Introduction to the fundamental aspects of common diseases affecting the brain including clinical features, animal models, genetics, neuropathology, synaptic function, and therapeutic targets. Prerequisite: NEUR 524.

NEUR 542 Hearing and Communication Neuroscience (4, Sp) (Enroll in BISC 521)

NEUR 599 Special Topics (2-4, max 8) Special topics providing background for instruction and research in neuroscience through lectures, discussions, assigned readings, and student presentations.

NEUR 790 Research (1-12, FaSpSm) Research leading to the doctorate. Maximum units which may be applied to the degree to be determined by the department. Graded CR/NC. Open only to neuroscience graduate students.

NEUR 794abcdz Doctoral Dissertation (2-2-2-2-0) Credit on acceptance of dissertation. Graded IP/CR/NC. Open only to neuroscience graduate students.